13  Hemolytic Anemia: Membrane Abnormalities & Enzymopathies

Key Points
  • Nonimmune hemolysis → consider hereditary hemolytic anemia; RBC morphology guides diagnosis
  • HS: most common congenital hemolytic anemia (Northern European, ~1:2,000)
    • Defects in ANK1, SLC4A1, SPTA1, EPB42 → ↓ surface-to-volume ratio
  • HE: RBC cytoskeleton protein defects (horizontal interactions); qualitative spectrin abnormalities common
  • Genetic testing recommended pre-splenectomy (efficacy & risks vary)
  • G6PD deficiency: episodic hemolysis w/ oxidative exposures; enzyme level falsely normal during acute episode
  • Glycolytic ATP maintains RBC membrane integrity; NADPH via HMP shunt maintains antioxidant capacity
  • Management: supportive care, folic acid, transfusions, iron chelation, splenectomy consideration
  • Mitapivat: oral PK activator available for PK deficiency

13.1 Introduction

  • Hemolysis: ↑ RBC destruction w/ compensatory ↑ erythropoiesis
  • Intrinsic causes: hemoglobin structure, RBC metabolism, membrane structure/permeability
  • RBC membrane structure
    • Lipid bilayer w/ transmembrane protein complexes
    • α & β-spectrin hexagonal lattice
    • Transverse & junction complexes (actin, protein 4.1)
    • Unique flexibility & deformability under stress
  • Clinical spectrum: asymptomatic (compensated) → symptomatic chronic hemolysis w/ transfusion dependence
  • Diagnosis critical: monitoring, supportive care, splenectomy efficacy/risks, targeted therapy availability vary

13.2 Monitoring Recommendations

Monitoring in RBC membrane & enzyme disorders
Parameter Frequency
Growth & development Baseline; q3-6 months year 1; then annual
CBC, retic, RBC indices Annual; ↑ frequency w/ accelerated hemolysis
Hemolysis markers (bili, LDH, hapto) q3-6 months if transfused; annual if not
Ferritin, TSAT, MRI T2* iron q3-6 months if transfused; annual if not
Bone health (25-OH vitamin D) Annual; consider late adolescence baseline
DEXA scan Consider late adolescence baseline
Echocardiogram Annual if regularly transfused or ↑ iron overload
Viral screening (HIV, HBV, HCV) Annual if regularly transfused
Abdominal ultrasound q2-3 years for gallstone disease (esp w/ Gilbert syndrome)
Echocardiogram Age >50 yr, pre-pregnancy, or pulmonary HTN concern

13.3 Red Cell Membrane Disorders

13.3.1 Hereditary Spherocytosis

Pathophysiology - Prevalence: most common congenital hemolytic anemia in Northern European descent (~1:2,000) - Defect: vertical interactions (membrane ↔︎ cytoskeleton) - ANK1 (ankyrin), SLC4A1 (band 3), SPTA1 (α-spectrin), EPB42 (protein 4.2) - ↓ membrane lipid & surface area via microvesiculation - Pathophysiology - ↓ surface-to-volume ratio → limited deformability - Splenic conditioning → progressive spherocytic changes → hemolysis - RBCs damaged traversing capillaries

Laboratory Evaluation - EMA binding assay: preferred test for membranopathies - Better sensitivity & specificity than osmotic fragility - Widely available; comparable to ektacytometry - Osmotic fragility test: 24-hr incubation at 37°C enhances sensitivity (inferior to EMA) - Osmotic gradient ektacytometry: laser diffraction; measures RBC deformability vs osmolality - Differentiates HS from HE & RBC cation permeability defects - Genetics: use w/ specialized RBC testing, especially no family history - Coinheritance of >1 variant common - Recommended pre-splenectomy (efficacy/risk differences) - RBC morphology: spherocytes on smear, ↑ MCHC

Severity classification in HS
Severity Hgb (g/dL) Retic (%) Bili (mg/dL) Features
Mild 11–15 3–8 1–2 Asymptomatic or few symptoms
Moderate 8–13 8–10 2–3 Mild-moderate; rare transfusion
Severe 6–8 >10 >3 Regular transfusions

Clinical Manifestations - Spectrum: asymptomatic (compensated) → severe hemolytic anemia w/ jaundice & splenomegaly - Complications - Gallstones: common in childhood/adolescence - Pigmented stones, early satiety, LUQ pain - Risk ↑ w/ Gilbert syndrome - Aplastic crisis: parvovirus → selective RBC suppression, reticulocytopenia - Megaloblastic anemia: rare; 2° to acquired folate deficiency (pregnancy, high demand) - Gilbert syndrome: present in majority of HS patients; ↑ unconjugated hyperbilirubinemia

Management - Folic acid supplementation: especially during ↑ erythropoiesis (pregnancy) or limited intake - Supportive care: monitoring, patient education on crisis signs/symptoms - Splenectomy: markedly improves hemolysis in almost all HS - Does NOT correct underlying RBC defect - Rule out RBC cation permeability defects pre-op (hemolysis risk if severe) - Severe/autosomal recessive HS: partial splenectomy data limited - Vaccinations: consider pre-splenectomy (encapsulated organisms)

13.3.2 Hereditary Elliptocytosis

Pathophysiology - Defect: RBC cytoskeleton protein interactions (horizontal interactions) - Qualitative spectrin abnormalities most common - SPTA1, SLC4A1, others - Coinheritance: >1 RBC gene variant not uncommon - RBC morphology: elliptocytes, may see mushroom-shaped RBCs (protein-specific)

Laboratory Evaluation - RBC morphology: guides initial diagnosis of hereditary hemolytic anemia - Specialized testing: morphologic abnormalities specific for causative protein defect - Note: Enzymopathies show nonspecific RBC morphology → differential w/ nonspherocytic hemolytic anemia

Clinical Manifestations & Management - Spectrum: variable severity; some w/ severe hemolysis requiring repeated transfusions - Splenectomy: markedly diminishes hemolysis but does NOT fully correct anemia in most

13.4 Red Cell Enzymopathies

13.4.1 Glycolytic Pathway & ATP Production

  • ATP generation: maintains RBC membrane integrity & oxygen affinity
  • Hexose monophosphate (HMP) shunt: produces NADPH
    • Maintains antioxidative capacity via glutathione system
    • Protects vs oxidative damage

13.4.2 Glucose-6-Phosphate Dehydrogenase (G6PD) Deficiency

Overview - Prevalence: most common enzymatic RBC disorder globally - Genetics: X-linked; G6PD synthesized 4-fold excess vs β-spectrin - Clinical spectrum: asymptomatic → hemolytic crises w/ oxidative stress

Clinical Manifestations - Asymptomatic until oxidative exposures occur - Hemolytic episodes: dark/cola-colored urine, jaundice - Triggers: oxidative stressors - Medications: antimalarials, sulfonamides, aspirin, NSAIDs, dapsone, nitrofurantoin - Infections: bacterial & viral (can precipitate crises) - Dietary: fava beans - Lab note: enzyme level may be falsely normal during acute episode - Test during non-hemolytic periods for accurate diagnosis

Management - Supportive: avoidance of trigger factors; transfusions if needed acutely - Education: medication triggers, dietary precautions critical - Monitoring: complications, folic acid supplementation - Family counseling: X-linked inheritance pattern

13.4.3 Pyruvate Kinase Deficiency

Pathophysiology - Enzyme role: glycolytic pathway ATP generation (essential) - Genetics: PKLR mutations → ↓ enzyme activity & ↓ ATP - Result: ↓ RBC membrane deformability → hemolysis

Management - Supportive: folic acid, transfusions if needed - Targeted therapy: mitapivat (oral PK activator) - Available for adults w/ PK deficiency - Advances beyond supportive care alone

13.5 Laboratory Evaluation Strategy

Initial workup: extravascular hemolysis pattern - ↑ LDH, ↑ indirect bilirubin, reticulocytosis, ↓ haptoglobin - Family history, RBC morphology review essential

Specialized testing order: 1. EMA binding assay: preferred for membranopathies - Sensitivity & specificity superior to osmotic fragility - Widely available 2. Osmotic gradient ektacytometry: if EMA unavailable - Differentiates HS from HE & cation permeability defects 3. Osmotic fragility test: 24-hr incubation at 37°C (enhanced sensitivity) 4. Genetic testing: w/ specialized RBC testing - Particularly if no family history of HS - Recommended pre-splenectomy

Sample handling - Transport quickly to lab w/ hot packs - Keep warm (cooling → agglutination & failed analysis) - Use Styrofoam container w/ hot packs

Clinical Pearl: Splenectomy in Hereditary Spherocytosis

Splenectomy improves hemolysis in nearly all HS patients, but does NOT correct underlying RBC defect. Rule out RBC cation permeability defects pre-op due to hemolysis risk in severe cases.

Clinical Pearl: G6PD Enzyme Testing Timing

Quantitative G6PD measurement may be falsely normal during acute hemolytic episode. Test during non-hemolytic periods or use alternative diagnostics when clinical suspicion high.

Clinical Pearl: Pyruvate Kinase Deficiency Treatment

Mitapivat (oral PK activator) now available for adults w/ PK deficiency & represents targeted therapy beyond supportive care.

13.6 Key Genes in RBC Membrane & Enzyme Disorders

Key genes in RBC membrane & enzyme disorders
Gene Protein Function Disorder
ANK1 Ankyrin-1 Vertical interactions; membrane-cytoskeleton linkage HS
SLC4A1 Band 3 (Cl−/HCO3− exchanger) Vertical & horizontal interactions HS, some HE
SPTA1 α-Spectrin Horizontal interactions; RBC skeleton HS, HE
EPB42 Protein 4.2 Membrane scaffolding; ankyrin binding HS
PKLR Pyruvate kinase Glycolytic ATP production PK deficiency
G6PD Glucose-6-phosphate dehydrogenase Pentose phosphate pathway; antioxidant G6PD deficiency